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Introduction
Part I: Life, meaning, and information
Exploring the philosophical background and scientific foundations of naturalist approaches to meaning and symbolism
Life sciences and the natural history of signs: can the origin of life processes coincide with the emergence of semiosis?
A proposal for a biosemiotic approach to digitalization: literacy as modelling competence
Threshold, meaning and life
How information gets its meaning
Part II: Semiosis and evolution
Inclusive Fitness teleology and Darwinian explanatory pluralism: a theoretical sketch and an application to current controversies
The origins and evolution of design: a stage-based model
Biosemiotics and applied evolutionary epistemology: a comparison
Extended synthesis and Jablonka and Lambs four-dimensional view of evolution
Part III: Physics, medicine, and bioenergetics
Physical intentionality: the phenomenological roots of biosemiotics
Cancer and cell death: a biosemiotic perspective
Biosemiotics and bioenergetics: two perspectives compared.

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